Designed, modeled, shaded/textured, lit, rendered and composited by Tosh Suo. Modeled in Zbrush + Maya. Rendered with Vray. Composited in Photoshop.
Featured in the 3dtotal cartoon gallery:
http://www.3dtotal.com/galleries/image/Cartoon/7610-sharks-by-tosh-suo-shark-underwater-zbrush-maya-vray-character-stylized